Micronutrients: Complete Guide to Vitamins

Comprehensive tutorial on all 13 essential vitamins - fat-soluble (A, D, E, K) and water-soluble (B complex, C). Functions, food sources, RDA, deficiency, toxicity, and clinical significance. From the NIH, IOM, and CDC.

This content is for informational purposes only. Always consult a healthcare professional.

Overview

Vitamins are organic compounds required in small amounts for normal physiological function. The human body cannot synthesize most vitamins in adequate amounts, making them essential dietary components. There are 13 essential vitamins classified into two groups based on solubility.

Property Fat-Soluble Vitamins Water-Soluble Vitamins
Vitamins covered A, D, E, K B complex (8) + C
Solubility Lipids and organic solvents Water
Absorption Requires bile and dietary fat Direct absorption
Transport Chylomicrons, carrier proteins Free in bloodstream
Storage Adipose tissue, liver (extensive) Limited (except B₁₂)
Excretion Bile (not urine) Urine (when intake exceeds needs)
Toxicity risk Higher (hypervitaminosis, especially A and D) Lower (excreted, except B₆ neuropathy and B₃ niacin flush)
Frequency of intake Dietary regularity over days/weeks Regular intake (days) needed

Fat-Soluble Vitamins

Vitamin A (Retinoids and Carotenoids)

Aspect Detail
Active forms Retinol, retinal (retinaldehyde), retinoic acid
Provitamins Beta-carotene, alpha-carotene, beta-cryptoxanthin (converted to retinol in body)
RDA (adults) Men: 900 μg RAE/day; Women: 700 μg RAE/day
Tolerable Upper Limit 3,000 μg RAE/day (preformed vitamin A)
Pregnancy 770 μg RAE/day; upper limit 3,000 μg (teratogenic above this)

Functions:

Function Mechanism
Vision (rhodopsin) 11-cis-retinal + opsin → rhodopsin (dark adaptation, color vision)
Gene expression Retinoic acid receptors (RAR, RXR) regulate transcription
Immune function Maintains mucosal barriers, T-cell differentiation
Epithelial integrity Differentiation of epithelial cells (skin, mucous membranes)
Reproduction Spermatogenesis, fetal development
Growth and development Bone remodeling, cell differentiation

Food Sources:

Food Serving Vitamin A (μg RAE) % DV
Beef liver (cooked) 85 g (3 oz) 6,582 731%
Sweet potato (baked) 1 medium (114 g) 1,403 156%
Carrot (raw) 1 medium (61 g) 509 57%
Pumpkin (canned) ½ cup (122 g) 953 106%
Spinach (cooked) ½ cup (95 g) 573 64%
Kale (cooked) ½ cup (65 g) 443 49%
Butternut squash (cooked) ½ cup (102 g) 572 64%
Cantaloupe ½ cup (80 g) 135 15%
Mango 1 medium (200 g) 112 12%
Red bell pepper 1 medium (120 g) 128 14%
Egg (whole, large) 1 (50 g) 75 8%

Deficiency:

Condition Description Populations at Risk
Night blindness (nyctalopia) Impaired dark adaptation (earliest sign) Malnutrition, fat malabsorption
Xerophthalmia Dry conjunctiva, Bitot spots, corneal keratomalacia Severe deficiency (leading cause of preventable blindness in developing countries)
Follicular hyperkeratosis Rough, dry skin (excess keratin) Chronic deficiency
Impaired immunity Increased infection risk (especially measles) Children in developing world
Increased mortality Measles, diarrhea, malaria Pre-school children

Toxicity (Hypervitaminosis A):

Type Cause Symptoms
Acute Single massive dose (≥25,000 μg RAE or ≥100,000 IU) Nausea, vomiting, headache, blurred vision, muscle incoordination
Chronic Daily intake >10,000 μg RAE (>30,000 IU) for months Dry skin, hair loss, joint pain, hepatotoxicity, pseudotumor cerebri
Teratogenicity Excess during pregnancy (≥3,000 μg RAE/day) Congenital malformations (fetal retinoid syndrome)
Carotenodermia High beta-carotene intake Yellow-orange skin (harmless, reversible)

Vitamin D (Calciferol)

Aspect Detail
Active form 1,25-dihydroxyvitamin D (calcitriol)
Precursor 7-dehydrocholesterol (skin) → cholecalciferol (D₃); ergocalciferol (D₂ from plants)
RDA (adults 19-70) 600 IU/day (15 μg/day)
RDA (adults >70) 800 IU/day (20 μg/day)
Tolerable Upper Limit 4,000 IU/day (100 μg/day)
Endogenous synthesis Sunlight (UVB, 290-315 nm) on skin — 10-30 min midday, 2-3x/week

Functions:

Function Mechanism
Calcium absorption Upregulates TRPV6 (intestinal calcium channel) and calbindin
Phosphate absorption Intestinal sodium-phosphate cotransporter
Bone mineralization Regulates calcium/phosphate deposition
Bone remodeling RANKL/RANK pathway (osteoclast differentiation)
Immune modulation Antimicrobial peptides (cathelicidin), T-cell regulation
Cell differentiation Antiproliferative effects (cancer prevention, observational)

Food Sources:

Food Serving Vitamin D (IU) % DV
Cod liver oil 1 tbsp (15 mL) 1,360 340%
Salmon (sockeye, cooked) 85 g (3 oz) 570 71%
Trout (rainbow, cooked) 85 g (3 oz) 540 68%
Tuna (canned, light) 85 g (3 oz) 40 5%
Sardines (canned) 2 sardines 47 6%
Milk (fortified) 1 cup (240 mL) 120 15%
Yogurt (fortified) 6 oz (170 g) 80 10%
Orange juice (fortified) 1 cup (240 mL) 100 13%
Mushrooms (UV-exposed) ½ cup (70 g) 366 46%
Egg yolk 1 large (17 g yolk) 37 5%
Beef liver (cooked) 85 g (3 oz) 42 5%

Deficiency:

Condition Description
Rickets (children) Soft, weak bones → bowing legs, delayed growth, bone pain
Osteomalacia (adults) Bone softening → diffuse bone pain, muscle weakness, fractures
Osteoporosis Reduced bone mineral density (long-term insufficiency)
Falls (elderly) Muscle weakness (proximal myopathy)
Impaired immune function Increased respiratory infections

Risk Groups for Deficiency:

Group Reason
Breastfed infants (unsupplemented) Low vitamin D in breast milk
Older adults ↓ Skin synthesis efficiency, ↓ sun exposure
Dark skin (high melanin) ↓ UVB absorption for vitamin D synthesis
Northern latitudes (>35°N) Limited UVB in winter months
Obese individuals Sequestration in adipose tissue
Malabsorption syndromes Crohn’s, celiac, cystic fibrosis, short bowel
Sun avoidance / covered clothing Inadequate UV exposure

Vitamin E (Tocopherols and Tocotrienols)

Aspect Detail
Active form α-Tocopherol (most bioactive)
Other forms β, γ, δ tocopherols + tocotrienols (lower bioactivity)
RDA (adults) 15 mg/day (22.4 IU)
Tolerable Upper Limit 1,000 mg/day (1,500 IU)
Primary function Lipid-soluble antioxidant (chain-breaking, peroxyl radical scavenger)

Functions:

Function Mechanism
Antioxidant Protects PUFAs in cell membranes from lipid peroxidation
Immune function Enhances T-cell mediated immunity
Cell signaling Inhibition of protein kinase C
Platelet aggregation Inhibition (vitamin E at high doses)
Neurological protection Myelin maintenance, nerve conduction

Food Sources:

Food Serving Vitamin E (mg α-TE) % DV
Wheat germ oil 1 tbsp (14 g) 20.3 135%
Sunflower seeds 30 g (¼ cup) 9.3 62%
Almonds 30 g (23 nuts) 7.3 49%
Hazelnuts 30 g (21 nuts) 4.3 29%
Sunflower oil 1 tbsp (14 g) 5.6 37%
Safflower oil 1 tbsp (14 g) 4.6 31%
Peanuts 30 g (28 nuts) 2.2 15%
Spinach (cooked) ½ cup (95 g) 1.9 13%
Avocado ½ fruit (100 g) 2.1 14%
Mango 1 medium (200 g) 1.5 10%

Deficiency:

Condition Description Context
Peripheral neuropathy Axonal degeneration (spinocerebellar tract) Fat malabsorption, genetic (abetalipoproteinemia)
Ataxia Loss of coordination Severe deficiency
Hemolytic anemia Red blood cell fragility Preterm infants
Myopathy Muscle weakness Chronic deficiency

Note: Vitamin E deficiency is extremely rare in healthy individuals. It occurs primarily with severe fat malabsorption (cystic fibrosis, Crohn’s, short bowel syndrome) or genetic defects in α-TTP (ataxia with vitamin E deficiency, AVED).

Vitamin K (Phylloquinone and Menaquinones)

Aspect Detail
Vitamin K₁ (phylloquinone) Plant source, primarily in green leafy vegetables
Vitamin K₂ (menaquinones, MK-4 to MK-13) Animal sources, bacterial production (gut microbiome)
Vitamin K₃ (menadione) Synthetic form (no longer used in human supplements)
AI (adequate intake, adults) Men: 120 μg/day; Women: 90 μg/day
No established UL Low toxicity potential

Functions:

Function Mechanism Proteins Activated
Blood coagulation γ-Carboxylation of clotting factors Factors II (prothrombin), VII, IX, X; Proteins C, S, Z
Bone metabolism Osteocalcin carboxylation (binds calcium to hydroxyapatite) Osteocalcin
Vascular health Matrix Gla protein (inhibits vascular calcification) Matrix Gla protein (MGP)
Kidney health Inhibition of nephrocalcinosis Nephrocalcinosis inhibitor

Food Sources (K₁):

Food Serving Vitamin K (μg) % DV
Kale (cooked) ½ cup (65 g) 531 443%
Spinach (cooked) ½ cup (95 g) 444 370%
Collard greens (cooked) ½ cup (75 g) 418 348%
Swiss chard (cooked) ½ cup (88 g) 286 238%
Broccoli (cooked) ½ cup (78 g) 110 92%
Brussels sprouts (cooked) ½ cup (78 g) 109 91%
Asparagus (cooked) ½ cup (90 g) 46 38%
Lettuce (romaine) 1 cup (47 g) 48 40%
Soybean oil 1 tbsp (14 g) 25 21%

Food Sources (K₂):

Food Serving K₂ Content
Natto (fermented soybeans) 100 g ~1,000 μg (MK-7)
Cheese (hard, aged) 50 g 5-50 μg (various MK)
Egg yolk 1 large 15-25 μg (MK-4)
Butter 14 g (1 tbsp) 2-5 μg (MK-4)
Chicken (dark meat) 100 g 10-15 μg (MK-4)
Sauerkraut 100 g 5-10 μg (MK-7)
Beef liver 100 g 5-10 μg (MK-4)

Deficiency:

Condition Context
Bleeding diathesis (elevated INR) Malabsorption, liver disease, antibiotic use (↓ gut bacteria)
Hemorrhagic disease of newborn Low placental transfer, low milk content, sterile gut
Warfarin overdose Antagonist of vitamin K cycle (intentional or inadvertent)
Osteoporosis (possible) Low vitamin K associated with low bone density (observational)

Water-Soluble Vitamins

Vitamin B₁ (Thiamin)

Aspect Detail
Active form Thiamin pyrophosphate (TPP)
RDA (adults) Men: 1.2 mg/day; Women: 1.1 mg/day
UL Not established (low toxicity; excess excreted)

Functions: Coenzyme in carbohydrate and amino acid metabolism (pyruvate dehydrogenase, α-ketoglutarate dehydrogenase, transketolase, branched-chain α-keto acid dehydrogenase). Nerve conduction (membrane ion channels).

Food Sources: Pork, whole grains, enriched grains, legumes, sunflower seeds, trout, tuna, macadamia nuts.

Deficiency (Beriberi):

Type Features
Wet (cardiac) High-output heart failure, peripheral vasodilation, edema
Dry (neurologic) Peripheral neuropathy (symmetrical), foot drop, wrist drop
Cerebral (Wernicke-Korsakoff) Confusion, ataxia, nystagmus (Wernicke); amnesia, confabulation (Korsakoff)
Gastrointestinal Anorexia, constipation, abdominal discomfort

Risk groups: Alcohol use disorder (most common in Western countries), chronic malnutrition, bariatric surgery, hyperemesis gravidarum, HIV/AIDS, dialysis.

Vitamin B₂ (Riboflavin)

Aspect Detail
Active forms Flavin mononucleotide (FMN), flavin adenine dinucleotide (FAD)
RDA (adults) Men: 1.3 mg/day; Women: 1.1 mg/day
UL Not established

Functions: Coenzyme for oxidation-reduction reactions (electron transport chain, fatty acid β-oxidation, branched-chain amino acid catabolism, vitamin B₆ activation).

Food Sources: Milk, yogurt, eggs, beef liver, almonds, mushrooms, spinach, enriched grains.

Deficiency (Ariboflavinosis): Cheilitis (angular stomatitis), glossitis (magenta tongue), seborrheic dermatitis, photophobia, corneal vascularization, normocytic anemia.

Vitamin B₃ (Niacin)

Aspect Detail
Active forms Nicotinamide adenine dinucleotide (NAD⁺, NADP⁺)
RDA (adults) Men: 16 mg NE/day; Women: 14 mg NE/day
UL 35 mg/day (for flushing form; nicotinamide has higher UL)
Note Can be synthesized from tryptophan (60 mg tryptophan ≈ 1 mg NE)

Functions: Over 400 enzymes require NAD/NADP — glycolysis, TCA cycle, fatty acid synthesis, DNA repair, PARP activity, sirtuin activation.

Food Sources: Chicken breast, beef liver, tuna, turkey, salmon, peanuts, brown rice, potatoes, green peas.

Deficiency (Pellagra):

Feature Description
Dermatitis Photosensitive rash (Casal necklace on neck), symmetrical
Diarrhea Inflammation of GI tract
Dementia Confusion, memory loss, depression, psychosis
Death If untreated (fourth D)

Vitamin B₅ (Pantothenic Acid)

Aspect Detail
Active form Coenzyme A (CoA)
AI (adults) 5 mg/day
UL Not established

Functions: CoA is central to fatty acid synthesis/oxidation, TCA cycle, acetylation reactions, cholesterol synthesis, steroid hormone formation, neurotransmitter synthesis.

Food Sources: Widespread — chicken liver, sunflower seeds, avocado, mushrooms, salmon, yogurt, potatoes, eggs, whole grains.

Deficiency: Extremely rare (“burning feet syndrome” — paresthesias). Seen only in severe generalized malnutrition.

Vitamin B₆ (Pyridoxine)

Aspect Detail
Active form Pyridoxal 5’-phosphate (PLP)
RDA (adults) 1.3 mg/day; >50 years: 1.7 mg (men), 1.5 mg (women)
UL 100 mg/day
Pregnancy 1.9 mg/day (used for nausea)

Functions: Coenzyme for >100 enzymes — amino acid metabolism (transamination, decarboxylation), heme synthesis (δ-aminolevulinic acid synthase), neurotransmitter synthesis (serotonin, dopamine, GABA, histamine), glycogen phosphorylase, steroid hormone regulation, homocysteine metabolism.

Food Sources: Chickpeas, beef liver, poultry, fish (tuna, salmon), potatoes, bananas, fortified cereals, sunflower seeds.

Deficiency: Microcytic hypochromic anemia (impaired heme synthesis), dermatitis, glossitis, depression, confusion, peripheral neuropathy. Impaired tryptophan conversion to niacin.

Toxicity: Peripheral neuropathy (sensory at doses >100 mg/day from supplements; reversible on discontinuation).

Vitamin B₇ (Biotin)

Aspect Detail
Active form Biotin (enzyme-bound)
AI (adults) 30 μg/day
UL Not established (no known toxicity)

Functions: Coenzyme for carboxylation reactions — acetyl-CoA carboxylase (fatty acid synthesis), propionyl-CoA carboxylase (odd-chain FA metabolism), pyruvate carboxylase (gluconeogenesis), β-methylcrotonyl-CoA carboxylase (BCAA metabolism).

Food Sources: Beef liver, eggs (yolk), salmon, avocado, pork, sweet potato, almonds, seeds, cauliflower.

Deficiency: Rare — dermatitis, alopecia, periorificial rash, neurological symptoms (depression, hallucinations), lactic acidosis. Risk factors: raw egg white consumption (avidin binds biotin), parenteral nutrition without biotin, prolonged anticonvulsant therapy, pregnancy (subclinical).

Vitamin B₉ (Folate)

Aspect Detail
Active form Tetrahydrofolate (THF)
RDA (adults) 400 μg DFE/day
Pregnancy 600 μg DFE/day (400 μg folic acid supplement recommended)
Lactation 500 μg DFE/day
UL (folic acid) 1,000 μg/day (from supplements/fortified foods only)

Functions: One-carbon transfer reactions — purine and pyrimidine (DNA/RNA) synthesis, amino acid metabolism (homocysteine → methionine), choline metabolism.

Food Sources:

Food Serving Folate (μg DFE) % DV
Beef liver (cooked) 85 g (3 oz) 215 54%
Lentils (cooked) ½ cup (99 g) 179 45%
Chickpeas (cooked) ½ cup (82 g) 141 35%
Asparagus (cooked) 4 spears (60 g) 89 22%
Spinach (cooked) ½ cup (95 g) 131 33%
Black-eyed peas (cooked) ½ cup (86 g) 105 26%
Avocado ½ fruit (100 g) 81 20%
Brussels sprouts (cooked) ½ cup (78 g) 78 20%
Fortified breakfast cereal 1 serving 100-400 25-100%

Deficiency:

Condition Description
Megaloblastic anemia Macrocytic RBCs, hypersegmented neutrophils, impaired DNA synthesis
Glossitis Smooth, red tongue
Neural tube defects Spina bifida, anencephaly (folate in periconceptional period prevents 50-70%)
Homocysteinemia Elevated homocysteine (potential CVD risk factor)
Cognitive decline Associated with low folate (elderly)

Toxicity: High-dose folic acid (≥1,000 μg/day from supplements) may mask B₁₂ deficiency, potentially worsening neurological damage.

Vitamin B₁₂ (Cobalamin)

Aspect Detail
Active forms Methylcobalamin, adenosylcobalamin
RDA (adults) 2.4 μg/day
Absorption Requires intrinsic factor (IF) from gastric parietal cells
Storage ~2-5 mg (liver); body stores last 3-5 years
UL Not established (extremely low toxicity)

Functions: Methionine synthase (homocysteine → methionine, requires methylcobalamin), methylmalonyl-CoA mutase (odd-chain FA → succinyl-CoA, requires adenosylcobalamin), myelin sheath maintenance.

Absorption:

Step Site Process
1. Protein release Stomach HCl + pepsin releases B₁₂ from food protein
2. Haptocorrin binding Stomach R-protein (haptocorrin) binds and protects B₁₂
3. Intrinsic factor Duodenum Pancreatic enzymes degrade haptocorrin; B₁₂ binds IF
4. IF-B₁₂ absorption Ileum (terminal) Cubam receptor (cubilin + amnionless) on enterocytes
5. Blood transport Portal circulation B₁₂ bound to transcobalamin II (holo-TC)

Food Sources (exclusive to animal products):

Food Serving Vitamin B₁₂ (μg) % DV
Beef liver (cooked) 85 g (3 oz) 71 2,958%
Clams (cooked) 85 g (3 oz) 84 3,500%
Sardines (canned) 85 g (3 oz) 8 333%
Salmon (cooked) 85 g (3 oz) 5 208%
Tuna (canned) 85 g (3 oz) 2.5 104%
Beef (ground, cooked) 85 g (3 oz) 2.5 104%
Milk 1 cup (240 mL) 1.2 50%
Yogurt (plain) 6 oz (170 g) 0.8 33%
Egg (hard-boiled) 1 large 0.6 25%
Nutritional yeast (fortified) 1 tbsp 2-4 83-167%

Deficiency:

Condition Description
Megaloblastic anemia Same as folate deficiency (macrocytic anemia)
Peripheral neuropathy Symmetrical paresthesias, numbness, loss of vibration sense
Subacute combined degeneration Dorsal columns (vibration/proprioception loss) + corticospinal tracts (spasticity, hyperreflexia)
Cognitive impairment Memory loss, confusion, dementia
Glossitis Smooth, red, painful tongue
Psychiatric Depression, paranoia, hallucinations

Risk Groups:

Group Mechanism
Vegans/vegetarians No dietary source (strict vegans need supplementation)
Older adults (≥50 years) 10-30% have atrophic gastritis → ↓ intrinsic factor, ↓ acid release
Pernicious anemia Autoimmune destruction of gastric parietal cells (autoantibodies to IF and H⁺/K⁺-ATPase)
Gastric bypass surgery ↓ IF-producing cells, altered anatomy
Crohn’s disease (ileal) Ileal inflammation reduces IF-B₁₂ absorption
Metformin use ↓ Calcium-dependent IF-B₁₂ uptake in ileum
PPI/H₂ blocker use ↓ Gastric acid reduces B₁₂ release from food

Vitamin C (Ascorbic Acid)

Aspect Detail
Active form L-Ascorbic acid (reduced form); dehydroascorbic acid (oxidized form)
RDA (adults) Men: 90 mg/day; Women: 75 mg/day
Smokers +35 mg/day (increased oxidative stress)
UL 2,000 mg/day
Functions Antioxidant, collagen synthesis, carnitine synthesis, neurotransmitter synthesis, iron absorption, immune function

Functions:

Function Mechanism
Collagen synthesis Hydroxylation of proline and lysine (vitamin C is essential cofactor for prolyl hydroxylase and lysyl hydroxylase)
Antioxidant Radical scavenger (regenerates vitamin E from tocopheroxyl radical)
Iron absorption Reduces Fe³⁺ to Fe²⁺ in stomach (enhances non-heme iron absorption)
Carnitine synthesis Co-factor for trimethyllysine and γ-butyrobetaine hydroxylase
Neurotransmitter synthesis Dopamine β-hydroxylase (norepinephrine), other neuropeptides
Immune function Stimulates leukocyte function (neutrophil chemotaxis, phagocytosis, apoptosis)
Epigenetic regulation Co-factor for TET enzymes and JmjC-containing histone demethylases

Food Sources:

Food Serving Vitamin C (mg) % DV
Red bell pepper (raw) ½ cup (75 g) 95 106%
Orange 1 medium (130 g) 70 78%
Kiwi 1 medium (69 g) 64 71%
Broccoli (cooked) ½ cup (78 g) 51 57%
Strawberries ½ cup (83 g) 49 54%
Brussels sprouts (cooked) ½ cup (78 g) 48 53%
Grapefruit juice ½ cup (120 mL) 47 52%
Green bell pepper (raw) ½ cup (75 g) 60 67%
Tomato juice ½ cup (120 mL) 22 24%
Mango 1 medium (200 g) 61 68%

Deficiency (Scurvy):

Feature Description
Gingival hyperplasia/bleeding Swollen, bleeding gums
Perifollicular hemorrhages Pinpoint hemorrhages around hair follicles
Ecchymoses Easy bruising
Impaired wound healing Poor collagen synthesis
Fatigue, malaise Early, non-specific symptom
Arthralgias Joint pain
Corkscrew hairs Fractured, coiled hairs
Anemia Possible contributory role (iron absorption, bleeding)

Risk Groups: Alcohol use disorder, smoking, restrictive diets, low fruit/vegetable intake, dialysis.

Toxicity: Generally well tolerated. GI distress (diarrhea, nausea) at >2,000 mg/day. Kidney stone risk (oxalate nephrolithiasis) — vitamin C is metabolized to oxalate; caution in stone formers.

B Vitamin Interactions

Interaction Mechanism Clinical Significance
Folate + B₁₂ Both involved in methionine synthase (homocysteine → methionine) Isolated B₁₂ deficiency treated with folic acid alone may worsen neurological damage
B₆ + B₁₂ + Folate Homocysteine metabolism (B₆: transsulfuration pathway; B₁₂, folate: remethylation) Combined may lower homocysteine (but mixed CVD trial results)
B₂ + B₆ FMN (B₂) required for PLP (B₆) formation B₂ deficiency can cause functional B₆ deficiency
B₃ (niacin) NAD/NADP essential for energy metabolism Interactions with multiple vitamin pathways

Key Takeaways

Vitamin Solubility Key Function Notable Deficiency Notable Food Source
A Fat-soluble Vision, immune, epithelial Night blindness, xerophthalmia Liver, sweet potato, carrots
D Fat-soluble Calcium/phosphate homeostasis Rickets, osteomalacia Sunlight, fatty fish, fortified milk
E Fat-soluble Antioxidant (membrane) Peripheral neuropathy (rare) Sunflower seeds, almonds
K Fat-soluble Coagulation, bone metabolism Bleeding (warfin-like) Kale, spinach, natto
B₁ (thiamin) Water-soluble Carb metabolism, nerve function Beriberi, Wernicke-Korsakoff Pork, whole grains, legumes
B₂ (riboflavin) Water-soluble Redox reactions Ariboflavinosis (cheilitis, glossitis) Milk, liver, eggs
B₃ (niacin) Water-soluble NAD/NADP synthesis Pellagra (dermatitis, diarrhea, dementia) Chicken, tuna, peanuts
B₅ (pantothenate) Water-soluble CoA synthesis Extremely rare Widespread in foods
B₆ (pyridoxine) Water-soluble Amino acid metabolism Microcytic anemia, dermatitis Chickpeas, liver, poultry
B₇ (biotin) Water-soluble Carboxylation reactions Rare (avidin-induced) Liver, eggs, avocado
B₉ (folate) Water-soluble DNA synthesis, methylation Megaloblastic anemia, NTDs Lentils, liver, greens
B₁₂ (cobalamin) Water-soluble Myelin, homocysteine metabolism Pernicious anemia, neuropathy Only animal foods (liver, clams)
C Water-soluble Collagen, antioxidant, iron absorption Scurvy Bell peppers, citrus, kiwi
  • The 13 essential vitamins are required for metabolism, growth, and disease prevention
  • Fat-soluble vitamins (A, D, E, K) require dietary fat for absorption and are stored in tissues (higher toxicity risk)
  • Water-soluble vitamins (B complex, C) require regular intake as excess is excreted (lower toxicity risk)
  • Vegans must supplement B₁₂ (not found in plant foods) and may need D, iodine, and iron
  • Alcohol use disorder is the leading cause of B₁ (thiamin) and B₃ (niacin) deficiency in Western countries
  • Pregnancy increases requirements for folate (neural tube prevention), iron, and vitamin D
  • Older adults frequently have impaired B₁₂ absorption and vitamin D synthesis